RE: bacterial pH

From: Howard Shapiro (hms@shapirolab.com)
Date: Tue Jun 27 2000 - 08:39:32 EST


Gerhard Nebe-von Caron notes, re Elaine Kunze's question about pH
measurement in E. coli-

>Due to the dye transporter systems the only working system for me has been
>carboxy-fluorescein-diacetate-succinimidylester. However in the absence of
>dual
>excitation ratio (as described in Howard Shapiro's book) your measurements are
>severely limited due to variations in the dye loading and cell volume => width
>of the distribution. Thus you only get a rough estimate. Unfortunately you can
>also not work in the exponential growth phase as that won't allow you dye
>loading in the first place. All a bit limited, the whole method unless you
>have
>a non-pumping strain..

I agree with Gerhard that the dye loading is the major problem, especially
with Gram-negative organisms such as E. coli.

A recent paper (Chitarra LG, Breeuwer P, van den Bulk RW, Abee T: Rapid
fluorescence assessment of intracellular pH as a viability indicator of
Clavibacter michiganensis subsp. michiganensis.  J Appl Microbiol 2000;
88:809-816) describes ratiometric flow cytometry of pH in a Gram-positive
organism using carboxyfluorescein succinimidyl ester (CFSE), taking the
ratio of emissions in the 520-540 nm and 560-590 nm range.  Even with the
ratiometric method, the "pH" curves observed were relatively broad;
however, while they did not completely discriminate heat-killed (nonviable)
and untreated (viable) cells, they did provide a usable, if crude,
indication of pH.  This area needs some more work.

-Howard



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