Re: Counting bacteria?

From: Howard Shapiro (hms@shapirolab.com)
Date: Sun May 16 1999 - 23:52:40 EST


Dr. Khalid Al-Hussein asks:

>     I have a graduate student who is interested in counting and 
>     determining the viability of bacteria (pesudomonas. sp, cultured in 
>     different pH concentrations) using Flow.
>     
>     Can anyone suggest how we could do this?.
>     

I would highly recommend the recent article on bacterial viability in
Current Protocols in Cytometry (Davey HM, Weichart DH, Kell DB, Kaprelyants
AS: Estimation of microbial viability using flow cytometry.  In: Robinson JP
et al (Eds): Current Protocols in Cytometry.  Wiley-Liss, New York,  1999.
11.3.1-11.3.20).

In general, there are two approaches to measurement of bacterial "viability"
using flow.  One examines the integrity of the cytoplasmic membrane; cells
which allow propidium, TO-PRO-3, Sytox Green (the last two are from
Molecular Probes) to enter, or which do not retain fluorescein or related
compounds produced by intracellular hydrolysis of esters such as fluorescein
diacetate, are considered to be nonviable. The second uses the presence of
membrane potential as an indicator of bacterial viability, using lipophilic
cationic dyes such as cyanines and rhodamine 123, or lipophilic anionic
dyes, such as oxonols, as membrane potential indicators.

The outer membrane of Gram-negative bacteria blocks entry of lipophilic
compounds, and membrane potential cannot be estimated satisfactorily in most
Gram-negative organisms unless the outer membrane is permeabilized using an
agent such as EDTA.  My experience with Pseudomonas aeruginosa is that the
organisms do not survive very long (probably less than 30 minutes) after
permeabilization, making it essential to do the preparation of cells and the
flow cytometry on a strict time schedule.  If you use an indicator of
membrane integrity, such as propidium or Sytox Green, permeabilization is
not necessary; cells which take up these dyes are likely to be nonviable.
However, cultures may also contain nonviable cells which do not take up the
dyes.

The addition of fluorescent beads at known concentration to aliquots of
sample allows you to get absolute counts of bacteria in fluorescence flow
cytometers in much the same way that absolute T-cell counts are now
obtained; both B-D and Coulter sell bead kits for this purpose, but it is
also possible to make up your own dilutions of beads from other vendors and
count them with a haemocytometer or, better still, with an impedance-based
(Coulter) or optical cell counter.  However, there may be a problem with
Pseudomonas aeruginosa because its forward scatter signal amplitude is
substantially smaller than those of other common bacteria, making it
difficult to trigger on forward scatter signals.  This could lead to
underestimates of bacterial counts.

Molecular Probes sells dye kits for bacterial viability estimation by flow;
you might try one.  Some people have had better luck with these than others.
By correlating your flow results with plate counts for a few days' worth of
experiments, you should be able to determine whether the kit will work with
your organisms.

-Howard Shapiro
 



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