proliferation markers for lymphocytes

From: DARZYNKIEWICZ ZBIGNIEW (DARZYNK@nymc.edu)
Date: Fri Jan 07 2000 - 15:30:02 EST


	A very reliable method to distinguish proliferating from
nonproliferating lymphocytes is based on differential staining of RNA and
DNA (See the original paper, PNAS, 73: 2881-84, 1976). Nonstimulated
lymphocytes have minimal RNA content and upon mitogenic stimulation (G0 to
G1 transition) their RNA  increases by at least one order of magnitude,
reflecting an increase in number or ribosomes. When cells enter S and G2/M
phases their DNA also is increased. Differential staining of DNA vs. RNA,
thus, allows one to discriminate between Go vs G1vs S vs G2/M cells and
detect as few stimulated cells as one per 10E4. They are also detected
early, upon entrance G1 phase and prior to initiation of DNA replication.
	Unfortunately, the companies that sell reagents and flow cytometers
discourage the use of this simple, rapid and inexpensive staining (it takes
~ 1 min, the yearly supply of the fluorochromes is below $10.-). perhaps
since it is more profitable to sell mAbs that also can detect mitogenically
activated cells. It is alleged that one of the dyes that can be used for
differential staining of RNA and DNA (acridine orange, AO) produces
"irreversible contamination" of flow cytometers. This is, of course,
nonsense, and AO is no worse than other strongly fluorescing dyes e.g such
as rhodamine 123.. We (and many other other labs that I know)  routinelly
use AO  e.g. on FACScan, followed by immunocytochemical samples, with no
problems as long as a rinse is given in between the samples, with a diluted
solution of a bleacher (chlorox) and detergent. For differential staining of
DNA and RNA one can use either AO or a combination of Hoechst and pyronine Y
dyes. The methods are described in Meth. Cell Biol. Vol 41, 402-419, 1994,
and in Current Protocols in Cytometry. 
Zbigniew Darzynkiewicz



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