Summary of tips to prevent cell clumping.

From: Jeff Barry (JBarry@picr.man.ac.uk)
Date: Tue Oct 31 2000 - 06:45:26 EST


Dear fellow Flow-ers,

A while back I asked for advice about how to prevent cell clumping together
during an apoptosis (propidium Iodide) assay. I've put together a summary of
the advice given in the hope that it may be of value to others in the
future.
We tried altering our procedure and while we were clump free we were also
cell free but it is early days yet.
Thanks for your help.
Regards,
Jeff Barry
Paterson Institute, Manchester, UK
General tips:
Firstly, if they are adherent or even if they grow in suspension but in
clumps, make sure that they are in as good a single cell suspension that you
can make before fixation
A biodegradable anticoagulant named ACD-A at 0.6% in PBS prevents cell
clumping of mononuclear blood cells and some stromal cells.
Also it can be a good idea to filter the cells through a 60-70um mesh to get
rid of the larger lumps (although clearly this will lose cells).
Keep cells suspended in 1 to 5 % BSA/media helps prevent clumping.
Try adding some EDTA (0.02%)
Try adding DNase at 50U/mL in all preparation media.
If you are using an alcohol fixation, try to resuspend the cells in the last
remaining volume after centrifugation and then add the alcohol drop wise
with mixing.  Another problem we have run into is over trypsinization.  It
sounds as if you are getting free DNA into your prep which will string
everything together.  We have experienced this also. Final tip may be to
assure yourself that the cell number is kept below 1X10e6/ml.  The cells
tend to clump less for some reason.
Where visible clumps are seen in the sample tubes, our users use nylon mesh
or Falcon 2270 or 2235 tubes with the mesh in the caps to filter the
samples. If the cell type they are using tends to clump, Accutase sold by
Phoenix Flow Systems o San Diego, CA. USA prevents clumping. They also sell
Accumax, an enzyme based product used for creating single cell suspensions
from tissue samples.

Ethanol fixation step:
Inadequate ethanol fixation is a primary cause of clumpage - it's a good
idea to vortex while fixing (careful though if looking for apoptotic cells
as they could explode at this point!). Also make sure that the pellet is as
free as possible of residual PBS before adding the ethanol.
Use EDTA or Dnase:
Secondly you could try using either EDTA or a low level concentration of
DNase in your PI solution as it is DNA from blown-up cells that can cause
clumps as well.
Use Anti-clumping agents:
I have read about a firm that's called TCS CellWorks in the UK, that sells
Accumax. It is used to disaggregate clumpy cells and viability seems to be
very good after treatment. You can read about it on
WWW.innovativecelltech.com <http://WWW.innovativecelltech.com>  for
additional information. You can also contact the office manager at TCS
CellWorks Ltd :Fay Crook : Fay@tcsgroup.co.uk <mailto:Fay@tcsgroup.co.uk>
PAA sell a product called Accutase, that's for prevent cell clumping. I
never used it, so I can't tell you if this product works or not

Use Nuclei:
A variation on this assay utilizes hypotonic lysis to isolate individual
nuclei instead of analyzing whole cells.  It is described in:
J Immunol Meth, 139: 271-279, 1991.  This should eliminate cell/cell
binding.

Protocols:
Protocol 1: APOPTOSIS AND CELL CYCLE ANALYSIS:
Courtesy of Nigel Miller

1.	1 million cells in 200ul PBS
2.	Add with stirring 2ml ice cold 70% ethanol 30% PBS.
3.	Leave 30 mins. On ice.
4.	Centrifuge 2000 rpm 5 mins.
5.	Resuspend pellet in 800ul PBS. If cells are clumped pass through a
25 gauge needle.
6.	Add 100 ul Rnase (1mg/ml-boiled 10' to destroy DNAse).
7.	Add 80 ul Propidium Iodide (0.5mg/ml).
8.	Incubate at 37 deg C for 30 mins.
9.	Analyse using the FACSCalibr..
10.	Use MODFIT to calculate DNA.

Protocol 2:
Courtesy of Dennis Young
Fix cells by adding the 10^6 cells (or less) in 1 ml to 10 ml ice cold 70%
EtOH while vortexing.
Use detergents (0.1% Triton-X 100)
Use filters (20 - 75 microns should work)
Use 25 gauge needle to resuspend cells right before sampling (Lastly) Lyse
cells and measure nuclei instead.

Protocol/Advice 3:
Courtesy of Kent Claypool
If preparing a single cell suspension is producing aggregates remove the
cytoplasm through a pepsin digest (references below) and look at bare
nuclei. Also the PI for a sub-G1 peak does not always work. The example at
the below web site had cell death that didn't produce a sub-G1 peak until a
positive control with Fas ligand was used. Then produced a great profile.
Cells have to have Fas receptor to trigger this apoptotic pathway. Might
want to also look at gel electrophoresis for DNA laddering, Bcl-2 by western
and/or flow, as well as the 'TUNEL' assay which can be done with nuclei
also. http://sciencepark.mdanderson.org/flow/files/DNA_PI.html
<http://sciencepark.mdanderson.org/flow/files/DNA_PI.html>



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